Main Missions Walkthrough: All 31 Steps and Boss Levels

The main mission chain is the 31-step spine of Palworld: it runs from waking up on the beach to killing a level 80 dragon at the bottom of the World Tree, and it gates fast travel, the Palbox, the eight towers and the entire endgame region along the way. This walkthrough puts every step in order with what it actually asks of you, the boss level and weakness at each tower, the tech you should have unlocked by the time you knock, and the EXP each mission pays.

Main Story at a Glance

Two numbers up front decide how you should play the story. First, the chain is heavily back-loaded — the last nine missions carry 698,145 EXP, which is 86% of everything the main story pays, so the first half feels like it pays nothing because it genuinely does. Second, the whole chain adds up to 816,018 EXP, and if that were your only source you would finish the game at level 43 while the final boss sits at level 80. The story cannot level you. Alphas, dungeons and sub missions have to fill the gap, which is why the walkthrough below flags a gear check at every tower instead of just a level.

Chapter 1: The Tutorial Missions (1–12)

The opening twelve missions pay 348 EXP between them, which one alpha kill will beat later, so treat them as an unlock checklist rather than a grind. They exist to hand you the four things the rest of the game runs on: a workbench, a Pal Sphere, a Palbox with a worker on it, and your first partner skill. The only steps worth slowing down for are 6 and 7 — the sphere tech sits at technology level 2 alongside the Palbox, so spend those first two points there and nowhere else. Step 12 gives you Foxparks' Harness, a flamethrower strapped to a Pal, and it is a genuine weapon upgrade at that stage of the game.

Chapter 2: The Eight Tower Bosses (13–22)

This block is the game. Eight faction towers, one mission each, 117,525 EXP across the whole run, and three of the fights bolt an extra objective on before the boss door opens — Marcus wants you to talk to two desert locals first, Victor wants five Syndicate scientists dead, Saya wants five ninjas, and Auri makes you clear six area barriers. Missions 14 and 15 are pure conversation, so the real count here is eight fights. The difficulty curve is not smooth either: the jump from Bjorn at level 60 to Auri at level 68 is the single biggest step in the chain, and Auri's fight is also the one that expects a flying mount.

#	Mission	Boss	Boss Lv.	Weak to	EXP
13	The Rayne Syndicate	Zoe & Grizzbolt	10	Ground	198
14	The Key Sphere	— head back to the Small Settlement	—	—	288
15	Zoe Rayne	— return to the tower and talk to Zoe	—	—	288
16	Thou Shalt Not Harm Pals	Lily & Lyleen	20	Fire	1,155
17	Flawless Victory Fixation	Axel & Orserk	30	Ground, Ice	3,345
18	Ruler of the Sands	Marcus & Faleris	40	Water	7,302
19	Interspecies Experiments	Victor & Shadowbeak	50	Dragon	15,210
20	A Fine Night for a Drink	Saya & Selyne	55	Dragon, Dark	21,294
21	Bearer of All Burdens	Bjorn & Bastigor	60	Fire	28,899
22	Hope Springs Eternal	Auri & Shaolong	68	Ice, Electric	39,546

Where Every Tower Is and How Fat the Health Bar Is

Tower bosses do not use the Pal's normal health — every fight multiplies it, and the multiplier is not tied to the boss's level. Bjorn is the outlier: ×62 health on a level 60 Pal makes it a longer fight than Auri's ×50 at level 68, and it is the wall most players hit on their first run. Grizzbolt's ×12 at the other end is why the first tower feels like a tutorial. All eight sit on the Palpagos map, and the coordinates below drop you at the entrance rather than the region — the tower pins are on our interactive map's tower boss layer if you would rather click than type.

Gear Check Before Each Tower

Every tower has a piece of tech that lands just before it, and knowing which one turns "I need more levels" into "I need three technology points". The pattern repeats all the way up: a body armour unlocks two to four levels under the boss, and a weapon class unlocks right on top of it. Two of the towers also punish you for the climb rather than the fight — Mount Obsidian cooks you on the way to Axel and Feybreak freezes you on the way to Bjorn, so the resistance variant matters more there than raw defense. The rows below list the best unlock available at or below each boss's level; the full ladders live in the armor by tech level guide and the weapon upgrade path.

Tower boss	Boss Lv.	Best armour unlocked by then	Weapon class	Sphere tier
Zoe & Grizzbolt	10	Cloth Outfit — Lv. 4	Three Shot Bow — Lv. 10	Pal Sphere
Lily & Lyleen	20	Pelt Armor — Lv. 12	Crossbow — Lv. 13	Giga Sphere — Lv. 20
Axel & Orserk	30	Heat Resistant Metal Armor — Lv. 25	Handgun — Lv. 28	Hyper Sphere — Lv. 27
Marcus & Faleris	40	Heat Resistant Refined Metal Armor — Lv. 40	Double-Barreled Shotgun — Lv. 39	Ultra Sphere — Lv. 35
Victor & Shadowbeak	50	Cold Resistant Pal Metal Armor — Lv. 50	Assault Rifle — Lv. 45	Legendary Sphere — Lv. 44
Saya & Selyne	55	Lightweight Plasteel Armor — Lv. 55	Laser Rifle — Lv. 51	Legendary Sphere
Bjorn & Bastigor	60	Cold Resistant Plasteel Armor — Lv. 54	Guided Missile Launcher — Lv. 56	Legendary Sphere
Auri & Shaolong	68	Ancient Armor — Lv. 68	Charge Rifle — Lv. 65	Legendary Sphere

Chapter 3: The Endgame Missions (23–31)

After the last tower the story stops being a boss ladder and becomes a fetch-and-open sequence: hear the rest of the plot, prove you have all eight Key Spheres, summon a whale god to punch a hole in the map, then work down through the World Tree. This is also where the game finally pays you — the nine missions below are worth 698,145 EXP, and Awakening alone hands over 143,736, more than the whole tower run combined. Mission 26 is the only step in the entire main story that gives an actual item.

The Eight Key Spheres

Gathering the Key Spheres is a checkpoint rather than a chore — each one is handed over by a tower leader, so if you have cleared all eight towers you already have all eight spheres and the mission completes the moment it opens. They are named after the seven deadly sins plus Original Sin, and none of them can be crafted, bought or dropped. The only way this step blocks you is if you skipped a tower, in which case the missing sphere tells you exactly which leader you still owe a fight.

Key Sphere of PrideKey Sphere of GreedKey Sphere of EnvyKey Sphere of WrathKey Sphere of LustKey Sphere of GluttonyKey Sphere of SlothKey Sphere of Original Sin

Panthalus and the Echoing Flute

This is the step people stall on, because nothing tells you where to start. Talk to the Ancient Civilization Researcher on the Deserted Islet at roughly -172, 33, then go collect four Echobones off four giant skeletons scattered across the map. The flute needs one of each, not four of the same bone, and it builds at the humble Primitive Workbench for 300 work — no ancient station, no ingots. Panthalus itself is a ten-minute timed fight on a small island, it is Water so Electric shreds it, and it is the only boss in the game that takes 20% extra damage. The capture at the end cannot fail, so throw a basic Pal Sphere and keep the good ones.

Material	Qty	Where it sits
Marine Echobone	1	122, -525 — western Grasslands, on the skull in shallow water
Silent Echobone	1	356, -110 — near the Altar of the Windcaller, inside the skull
Seafoam Echobone	1	-519, -548 — Mount Obsidian, under the ribcage
Tidewind Echobone	1	-262, 485 — snowfields, on the frozen lake by the teleport
Total	4 bones, 300 work	Primitive Workbench → 1 Echoing Flute

Full attack list, the wind-column trick for the tidal wave and the drop table are in the Panthalus fight and World Tree entry guide.

Awakening: Zenara and Astralym

The last mission is not a tower duel, it plays like a raid. Astralym has no element at all, which means nothing on the element matchup chart gives you a bonus against it — the counter-picking habit you built over eight towers is worthless here. Its first-fight health multiplier is ×40, modest next to Bjorn's ×62, but the fight is long because you are doing flat damage with no type bonus. Two of its eight moves are Dragon, and Dragon beats Dark, so the Dark-heavy squad most players finish the game with is the exact roster that takes extra damage. Swap in something neutral to its kit before you walk in.

The team, the passive stack and the full move list are in the Zenara and Astralym fight guide.

Recommended Level for Every Story Fight

The game posts a suggested level at each tower gate, and it runs a few points above the boss's own level for the mid-game fights — the PIDF tower asks for 45 against a level 40 boss, and the Genetic Research Unit asks for 55 against a level 50 one. That gap is not padding: those two are the fights where the recommended level is really a gear-tier check in disguise. The line worth remembering is the last one. There is a 12-level jump between the final tower and the final boss, and no main mission in between pays enough EXP to cover it, so that gap gets closed by alphas, raids and dungeons or it does not get closed at all.

Fight	Boss Lv.	Recommended Lv.	HP ×
Zoe & Grizzbolt	10	10+	×12
Lily & Lyleen	20	20+	×25
Axel & Orserk	30	30+	×25
Marcus & Faleris	40	45+	×27
Victor & Shadowbeak	50	55+	×28
Saya & Selyne	55	55+	×54
Bjorn & Bastigor	60	65+	×62
Auri & Shaolong	68	70+	×50
Panthalus	—	70+	×42
Zenara & Astralym	80	80	×40

Story Order in One Paragraph

Do the twelve tutorial steps in whatever order the game throws them, spend your first technology points on the sphere and the Palbox, and get Foxparks' Harness before you leave the Grasslands. Then run the towers in order — Zoe, Lily, Axel, Marcus, Victor, Saya, Bjorn, Auri — clearing the extra objective attached to Marcus, Victor, Saya and Auri, and topping up your armour tier between each one rather than your level. Bjorn is the wall. After Auri, the eight Key Spheres are already in your bag, so it is straight into the Echobone hunt, the Echoing Flute, and a Panthalus fight that Electric Pals trivialise. Enter the World Tree, clear the three mid-bosses, drop into the abyss, and bring a squad with nothing Dark in it to Astralym.

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